The story takes place several years after the events of Devil May Cry 4. A man known as "V" comes to the Devil May Cry office to hire Dante to solve a series of demon attacks occurring across the world. Meanwhile, Nero has set up his own demon hunting agency based out of a van adorned with a neon "Devil May Cry" sign Dante gave him with support by his engineer Nico, who built him a robotic prosthetic arm called the "Devil Breaker" after his original Devil Bringer arm was taken by a mysterious hooded individual. Nero's hunt for the hooded figure, a powerful demon named Urizen, brings him to Red Grave City, which Dante deduces as the epicenter of the demon attacks. The city then becomes ground zero for the growth of the Qliphoth tree, as Urizen nourishes its fruit with human blood so he can consume it and become king of the Underworld.
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